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Billingham, Rupert Everett [Bill] (1921–2002), immunologist, was born on 15 October 1921 at 95 Portway, Warminster, Wiltshire, the only child of Albert Everett Billingham (1889–1977) and his wife, Helen Louise (1888–1978), daughter of Herbert Green. His father's family originated in the industrial Black Country...

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Burnet, Sir Frank Macfarlane (1899–1985), virologist and immunologist, was born on 3 September 1899 in Traralgon, Gippsland, Victoria, Australia, the second of seven children of Frank Burnet (b. 1856), a banker of Scottish descent, and Hadassah Pollock Mackay (b. 1872), the daughter of a schoolteacher who had also emigrated from ...

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Sir Frank Macfarlane Burnet (1899–1985) by Walter Bird, 1958 © National Portrait Gallery, London

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Milstein, César (1927–2002), molecular biologist and immunologist, was born on 8 October 1927 in Bahía Blanca, a provincial town in Argentina, the second of three sons of Lázaro Milstein, a salesman who had travelled to Argentina at the age of fourteen as a Jewish immigrant from the ...

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César Milstein (1927–2002) by Nick Sinclair, 1993 © Nick Sinclair; photograph National Portrait Gallery, London